This study reports on the first map-based cloning of an NBG gene on chromosome 4 denoted NBG4 which is a novel allele of Dwarf 11 encoding a cytochrome P450 (CYP724B1) involved in brassinosteroid (BR) biosynthesis. In addition to the dwarf and clustered panicle as previously reported in the allelic mutants nbg4 grains also displayed retarded germination and NBG due to the physical constraint of deformed hulls caused
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